Birding Top 500 Counter

Hybridization in Hummingbirds

The colored grids below depict hummingbird hybrid combinations that have been adequately documented in the United States, north of Mexico. Many more combinations have been documented with other species south of the border. The dark BROWN grids depict combinations described in the scientific literature (with initial or key citations listed in the grid). RED grids depict combinations documented via photograph or convincing write-ups by researchers, but not published in scientific journals. See the references below the table for information on these combinations, many of which are shown on websites (links provided).

The Rufous x Anna's combination apparently arises from early descriptions of Selasphorus floresii Gould (Simon 1910, Ridgway 1909). There is apparently much disagreement on the parentage of this "species" (Banks & Johnson 1961), hence it is shown in GREEN. Banks and Johnson (1961) also disputed earlier reports of Costa's x Black-chinned, but that combination has since been described from other specimens (Short & Phillips 1966).

SPECIES  Broad-
billed
Violet-
crowned
Berylline Blue-
throated
Magnificent Lucifer Ruby-
throated
Black-
chinned
Anna's Costa's Calliope Broad-
tailed
Rufous Allen's
Broad-
billed
  Bishop 1906     Short & Phillips 1966                  
Violet-
crowned
Bishop 1906                          
Berylline                            
Blue-
throated
                Baldridge, et. al. 1983          
Magnificent Short & Phillips 1966                          
Lucifer                            
Ruby-
throated
                         
Black-
chinned
                 Jeffries 1888
Banks & Johnson 1961
Short & Phillips 1966    Banks & Johnson 1961   Lynch & Ames 1970
Anna's       Baldridge, et. al. 1983        Jeffries 1888
Banks & Johnson 1961
   Wells, et. al. 1978 Berloiz 1930
Graves & Newfield 1996
 
Banks & Johnson 1961

Summarized by Banks & Johnson 1961
Costa's               Short & Phillips 1966  Wells, et. al. 1978   Hartert 1900
Banks & Johnson 1961
 
Huey 1944    
Calliope                 Berloiz 1930
Graves & Newfield 1996
 Hartert 1900
Banks & Johnson 1961
 
      Thayer & Bangs 1907  
Broad-
tailed
                Banks & Johnson 1961   Huey 1944        
Rufous                
Banks & Johnson 1961
 
   Thayer & Bangs 1907       Newfield 1983
Allen's               Lynch & Ames 1970
Summarized by Banks & Johnson 1961
       Newfield 1983  

Here we provide information for those interested in this fascinating ornithological feature. We encourage researchers who encounter suspected hybrids to carefully document their discovery and submit it to a refereed journal for publication. If you would like to share your photos with other ummingbird researchers through this page, or if you are aware of published literature we should add to our compendium below, please email me.

ONLINE PHOTOS / DOCUMENTATION OF HYBRIDS


BLACK-CHINNED x CALLIOPE

May 24, 2008
Garden Valley, Idaho
Fred Bassett

August 4, 2004
near Rockland, Idaho
Terry L. Peterson & Stacy J. Peterson

August 8, 2004
High Valley, Idaho
Peggy L. Peterson & Stacy J. Peterson

August 8, 2004
High Valley, Idaho
Greg D. Wardwell & Stacy J. Peterson

   

Comparative page
for three Black-chinned x Calliope (August 2004) above. Interestingly, within days of the August 2004 BCHU x CAHU birds above, Fred Bassett banded another similar one in Twin Falls County, Idaho. A central diagnostic tail feather was collected from that bird.


BLACK-CHINNED x RUBY-THROATED

April 30, 2005
Concan, Texas
Bob Sargent & Fred Bassett

   

BLACK-CHINNED x BROAD-TAILED

July 2007
Lakewood-Green Mtn, Colorado
Photographed by Mark Chavez

   

RUFOUS x CALLIOPE

May 9, 2005
Lincoln, Montana
Ned & Gigi Batchelder

April 20, 2005
Mariposa, California
Rita Colwell

 

RUFOUS x ANNA'S

April 22, 2008
Eugene, Oregon
Found by Maitreya & Barry McKenzie
Observed by Dave Irons & others
Photographed by Peter Patricelli
   

BROAD-TAILED x CALLIOPE

[Need date] 2003
Red Lodge, Montana
Ned & Gigi Batchelder

   

Weird Plumages

SY-Male (lack of molt) Broad-tailed Hummingbird
This is a (presumably) rare example of an "early-blooming gorget" combined with retained juvenile plumage! In other words, this is likely a funny-looking "second-year" (hatched in 2004) male Broad-tailed.

May 2005, near Rockland, Idaho
Stacy J. Peterson


TRADITIONAL PUBLICATIONS dealing with hybridization in hummingbirds (and selected articles dealing with avian taxonomic issues)

Articles from all but the last few years in the journals Wilson's Bulletin, Journal of Field Ornithology, Auk, Condor, North American Bird Bander, Studies in Avian Biology, and Pacific Coast Avifauna may be available as PDF files through SORA: Searchable Ornithological Research Archive. Search by the authors name and a keyword in the title. Journal of Experimental Biology articles may be found here.

  • Baldridge, F.A., L.F. Kiff, S.K. Baldridge, and R.B. Hansen. 1983. Hybridization of a Blue-throated Hummingbird in California. Western Birds 14:17-30.
  • Banks, R.C. and N.K. Johnson. 1961. A review of North American hybrid hummingbirds. Condor 63:3-28.
  • Berlioz, J. 1930. A new hummingbird hybrid, Calypte anna x Stellula calliope. Condor 32:215.
  • Binford, L.C. 1985. Re-evaluation of the "hybrid" hummingbird Cyanthus sordidus X C. latirostris from Mexico. Condor 87:148-150.
  • Bishop, L.B. 1906. Uranomitra salvini in Arizona. Auk 23:337-338. [Uranomitra salvini = Violet-crowned x Broad-billed hybrid; see Griscom 1934]
  • Bleiweiss, R., J.A.W. Kirsch, and J.C. Matheus. 1994. DNA-DNA hybridization evidence for subfamily structure among hummingbirds. Auk 111:8-19.
  • Bleiweiss, R., J.A.W. Kirsch, and J.C. Matheus. 1997. DNA hybridization evidence for the principal lineages of hummingbirds (Aves: Trochilidae) Mol. Biol. Evol. 14:325-343.
  • Cockrum, E.L. 1952. A check-list and bibliography of hybrid birds in North America north of Mexico. Wilson Bull. 64:140-159.
  • Elgar, R.J. 1977. The breeding of a hybrid hummingbird Brown Violetear X Sparkling Violetear. Avic. Mag. 83: 130-134.
  • Jefferies, J.A. 1888. A description of an apparently new species of Trochilus from California. Auk 5:168-169.
  • Gerwin, J. A. and R.M. Zink. 1998. Phylogenetic patterns in the Trochilidae. Auk 115:105-118.
  • Gill, F.B., F.J. Stokes and C. Stokes. 1973. Contact zones and hybridization in the Jamaican Hummingbird, Trochilus polytmus (L.). Condor 75:170-176.
  • Graves, G.R. 1990. Systematics of the "Green-throated Sunangels" (Aves: Trochilidae): valid taxa or hybrids? Proc. Biol. Soc. Wash. 103:6-25.
  • Graves, G.R. 1996. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 2. Hybrid origin of Eriocnemis soderstromi Butler. Proc. Biol. Soc. Wash. 109:764-769.
  • Graves, G.R. 1997. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 3. Parentage of Lesbia ortoni Lawrence. Proc. Biol. Soc. Wash.110:314-319.
  • Graves, G.R. 1997. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 4. Hybrid origin of Calothorax decoratus Gould. Proc. Biol. Soc. Wash. 110:320-325.
  • Graves, G.R. 1998. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 5. Probable hybrid origin of Amazilia distans Whetmore & Phelps. Proc. Biol. Soc. Wash. 111:28-34.
  • Graves, G.R. 1998. Diognoses of hybrid hummingbirds (Aves: Trochilidae). 6. An intergeneric hybrid, Aglaiocercus kingi x Metallura tyrianthina, from Venezuela. Proc. Biol. Soc. Wash. 111:511-520.
  • Graves, G.R. 1999. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 7. Probable parentage of Calliphlox iridescens Gould 1860. Proc. Biol. Soc. Wash. 112:443-450.
  • Graves. G.R. 1999. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 8. A provisional hypothesis for the hybrid origin of Zodalia glyceria (Gould, 1858). Proc. Biol. Soc. Wash. 112:491-502.
  • Graves, G.R. 2001. Diangoses of hybrid hummingbirds (Aves: Trochilidae). 9. Confirmation of the hybrid origin of Coeligena purpurea Gould, 1854. Proc. Biol. Soc. Wash. 114:42-50.
  • Graves. G.R. 2003. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 10. Cyanomyia salvini Brewster, 1893, is an intergeneric hybrid of Amazilia violiceps and Cynanthus latirostris. Proc. Biol. Soc. Wash. 116:293-300.
  • Graves, G.R. 2003. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 11. Documentation of an intergeneric woostar hybrid, Calliphlox mitchelli x Chaetocercus mulsant. Proc. Biol. Soc. Wash. 116:820-826.
  • Graves, G.R. 2003. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 12. Amazilia bangsi Ridgway, 1910, is an intragernic hybrid, Amazilia tzacatl x Amazilia rutila. Proc. Biol. Soc. Wash. 116:847-852.
  • Graves, G.R. and N.L. Newfield. 1996. Diagnoses of hybrid hummingbirds (Aves: Trochilidae). 1. Characterization of Calypte anna X Stellula calliope and the possible effects of egg volume on hybridization potential. Proc. Biol. Soc. Wash. 109:755-763.
  • Graves, G.R. and R.L. Zusi. 1990. An intergeneric hybrid hummingbird (Heliodoxa leadbeateri X Heliangelus amethysticollis) from northern Colombia. Condor 92:754-760.
  • Griscom, L. 1934. The ornithology of Guerrero, Mexico. Bull. Mus. Comp. Zool. 75:378. [Concludes Uranomitra salvini = Violet-crowned x Broad-billed hybrid]
  • Hartert, E.J. 1900. [Exhibition of hybrid hummingbirds.] Bull. Brit. Ornith. Club 10:39-40.
  • Hinkelmann, C. 1996. Evidence of natural hybridization in hermit hummingbirds (Phaethornis spp.) Bull. Br. Ornithol. Club 116:5-14.
  • Hinkelmann, C., B. Nicolai and R.W. Dickerman. 1991. Notes on the hitherto unknown specimen of Neolesbia nehrkorni (Berlepsch, 1887; Trochilidae) with a discussion of the hybrid origin of this 'species'. Bull. Br. Ornithol. Club 111:190-199.
  • Huey, L.M. 1944. A hybrid Costa's x Broad-tailed hummingbird. Auk 61:636-637.
  • Lynch, J.F. and P.L. Ames. 1970. A new hybrid hummingbird, Archilochus alexandri X Selasphorus sasin. Condor 72:209-212.
  • Newfield, N.L. 1983. Records of Allen's Hummingbird in Louisiana and possible Rufous X Allen's hummingbird hybrids. Condor 85:253-254.
  • Pierotti, R. and C.A. Annett. 1993. Hybridization and male parental investment in birds. Condor 95:670-679.
  • Ridgeway, R. 1909. Hybridism and generic characters in the Trochilidae. Auk 26:440-442.
  • Short, L.L. 1969. Taxonomic aspects of avian hybridization. Auk 86:84-105.
  • Short, L.L. and A.R. Phillips. 1966. More hybrid hummingbirds from the United States. Auk 83:253-265.
  • Sibley, C.G. 1957. The evolutionary and taxonomic significance of sexual dimorphism and hybridization in birds. Auk 59:166-191.
  • Taylor, W.P. 1909. An instance of hybridization in hummingbirds, with remarks on the weight of generic characters in the Trochilidae. Auk 26:291-293.
  • Thayer, J.E. and O. Bangs. 1907. Another hybrid hummingbird - Selasphorus rufous + Atthis calliope - from California. Auk 24:312-313.
  • Wells, S. and L.F. Baptista. 1979. Displays and morphology of an Anna's X Allen's hummingbird hybrid. Wilson Bull. 91:524-532.
  • Wells, S., R.A. Bradley and L.F. Baptista. 1978. Hybridization in Calypte hummingbirds. Auk 95:537-549.
  • Williamson, F.S.L. 1957. Hybrids of the Anna's and Allen's hummingbirds. Condor 59:118-123.
  • Zink, R.M. and M.C. McKitrick. 1995. The debate over species concepts and its implications for ornithology. Auk 112:701-719.


OTHER DOCUMENTED HYBRID REPORTS

Berylline x Magnificent
August 2000, Miller Canyon, Arizona
Photographed by Charles Melton, published in Birding (37:235)

Lucifer x Black-chinned
unspecified date and location
Photographed by Dale & Marian Zimmerman, published in Williamson, S.L. 2001. Field Guide to Hummingbirds of North America. Houghton Mifflin. Boston.


Home

Send questions or comments to: Stacy Jon Peterson